NPP = GPP - R

Net primary production (NPP) is the chemical energy store in plant biomass after respiratory losses to the environment have been taken into account
‘Net’ means ‘overall’ (i.e after any necessary deductions have been taken into account) so net primary production refers to the overall chemical energy available after the necessary deductions (in this case energy lost through respiration) have been taken into account. Again, make sure you are consistent with your units - if an exam question gives GPP and R in joules (per unit area), you should give the NPP in joules (per unit area) too, unless the question specifically asks otherwise.Note the difference between production and productivity! One is a rate, the other isn't.
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